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Seehausen, Ole; Koetsier, Egbert; Schneider, Maria Victoria; Chapman, Lauren J.; Chapman, Colin A.; Knight, Mairi E.; Turner, George F.; van Alphen, Jacques J. M.; Bills, Roger (2003). Nuclear markers reveal unexpected genetic variation and a Congolese-Nilotic origin of the Lake Victoria cichlid species flock. Proceedings of the Royal Society. Series B - biological sciences, 270(1511), pp. 129-137. Royal Society of London 10.1098/rspb.2002.2153